Zant

Zant is the leading villain throughout most of the game. He is a member of the Twili race and the self-proclaimed ruler of the Twilight. This garnered the name, Usurper King of the Twilight. Zant invades Hyrule Castle alongside several shadow beings and forces Princess Zelda to surrender. He appears sparingly throughout the game, confronting Midna and Link on several fronts. After gathering the three fused shadows, Zant steals them from Midna and puts the shadow crystal within Link. This makes Link remain in his wolf form even when in the world of light.

Zant later attempts to destroy the Mirror of Twilight, but is unable to do so, so instead, breaks it into pieces and scatters it throughout Hyrule. Before the battle with Zant, It is revealed that Zant was acting as a tool for his God, Ganondorf.